A newly created directory inherits permissions (rwx stuff) from their parent and owner/group from the creator. The Default_Directory_Attributes is only used for a directory that doesn't have any owner/permission information at all which means it was likely created outside of the server and doesn't have a .ioFTPD file in it.
Please note that the R attribute for directories acts like R and X as far as unix would be concerned. Use private dirs if you want weird behavior where you would only have one of those attributes. This frees up the X bit which was intended to be used in archiving scripts so you could lock dirs from being archived and/or auto-wiped for space.
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