Hmm.
According to
https://tools.ietf.org/html/rfc3659#section-4.2
Quote:
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Where the command is correctly parsed but the size is not available, perhaps because the pathname identifies no existing entity or because the entity named cannot be transferred in the current MODE and TYPE (or at all), then a 550 reply should be sent.
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Just in case, can you paste output of the FEAT command and also build of FlashFXP?