got it working setting those values (over my inital settings) in Vsftpd on the 2 VMs :
pasv_address=1.2.3.4
connect_from_port_20=NO
and
pasv_address=5.6.7.8
connect_from_port_20=NO
Also make sure both Vms firewalls allow connections and transfers with the other one.
Then it started to work whatever the settings in FlashFXP (SSL or not, PASV upload or not).
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